Syllabus Edition

First teaching 2015

Last exams 2025

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Species Richness & Diversity

Alistair Marjot

Written by: Alistair Marjot

Reviewed by: Bridgette Barrett

Species Richness & Diversity

What is species richness?

  • Species richness is the number of species in a community or defined area and can be a useful comparative measure in some cases

  • However, in other cases, species richness can be a misleading indicator of diversity as it does not take into account the number of individuals of each species

  • Once the abundance of each species in an area has been recorded, the results can be used to calculate the species diversity for that area

    • Species diversity looks at the number of different species in an area but also the evenness of abundance across the different species (i.e. their relative abundances)

Species Richness vs Diversity

  • An index of diversity is a measurement that describes the relationship between the number of species present and how each species contributes to the total number of organisms that are present in that community

    • It is a much more informative measurement than species richness and conservationists often favour the use of an index of diversity as it takes into account both species richness and evenness

    • A commonly used index of diversity is known as Simpson’s Diversity Index

Example

  • Area 1 and Area 2 both contain 4 tree species

  • However, Area 2 is actually dominated by one species and in fact, one of the species is very rare (only one individual)

  • Although the two areas have exactly the same species richness, Area 1 has a higher species evenness (and therefore a higher overall species diversity) than Area 2

  • This example illustrates the limitations of using just species richness on its own


Area 1 and Area 2 have the same species richness but different species evenness. As it has a higher species evenness, the overall species diversity of Area 1 is higher than that of Area 2, as species diversity takes both richness and evenness into account

Calculating Simpson’s Diversity Index

  • A group of students used the kick sampling technique to collect, identify and count the invertebrates inhabiting a river

  • Samples were obtained from different sites along the course of the river

  • The data was used to calculate the Simpson's Diversity Index at two different river sites

  • This index of diversity is useful when comparing two similar habitats, or the same habitat over time

  • The formula for calculating Simpson's Diversity Index, D, is:

Species

Mean number of organisms per m2 of river bed

Site A

Site B

Mite

14

0

Snail

9

0

Leech

3

26

Worm

0

6

Flat worm

132

9

Mayfly nymph

43

0

Olive mayfly nymph

154

0

Midge Larva

0

10

Blackfly larva

77

0

Caddis larva

15

1

Fish

1

0

Freshwater shrimp

211

6

Water hog louse

0

40

Site A

Species

Number (n) 

n (n-1)

Mite

14

182

Snail

9

72

Leech

3

6

Worm

0

0

Flat worm

132

17 292

Mayfly nymph

43

1 806

Olive mayfly nymph

154

23 562

Midge Larva

0

0

Blackfly larva

77

5 852

Caddis larva

15

210

Fish

1

0

Freshwater shrimp

211

44 310

Water hog louse

0

0

Total

N= ∑n= 659

∑n(n-1)= 93 292

bold italic D equals bold italic space fraction numerator N stretchy left parenthesis N minus 1 stretchy right parenthesis over denominator capital sigma n stretchy left parenthesis n minus 1 stretchy right parenthesis end fraction bold italic equals fraction numerator space 659 stretchy left parenthesis 658 stretchy right parenthesis over denominator 93292 end fraction bold italic space equals space 4.65

Site B

Species

Number (n) 

n (n-1)

Mite

0

0

Snail

0

0

Leech

6

30

Worm

26

650

Flat worm

9

72

Mayfly nymph

0

0

Olive mayfly nymph

0

0

Midge Larva

10

90

Blackfly larva

0

0

Caddis larva

1

0

Fish

0

0

Freshwater shrimp

6

30

Water hog louse

40

1 560

Total

N= ∑n= 98

∑n(n-1)= 2 432



bold italic D equals bold italic space fraction numerator N stretchy left parenthesis N minus 1 stretchy right parenthesis over denominator capital sigma n stretchy left parenthesis n minus 1 stretchy right parenthesis end fraction bold italic equals fraction numerator space 98 stretchy left parenthesis 97 stretchy right parenthesis over denominator 2432 end fraction bold italic space equals space 3.91

  • Site A was located just 5 km downstream from the river’s source

  • Site B was located 50 m downstream from a sewage inlet pipe

  • The lower diversity index for site B reflects the stress placed upon the river as a consequence of the pollution load from the sewage inlet point; although the river is recovering from the pollution, sensitive species are unable to tolerate the unfavourable abiotic conditions and species diversity is reduced

Examiner Tips and Tricks

Remember, this index of diversity is only useful when comparing two similar habitats, or the same habitat over time.

You will be provided with the formula for Simpson’s Index in the exam but you need to know how to use it to calculate Simpson’s Diversity Index for example sets of data.

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